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Published 2014
Doak (1978) has described these fish as ‘swift and wary herbivores [which] more resemble deer than grazers such as sheep’. The three cusps (whence tricuspidata) on their close-set teeth enable them to crop the seaweeds which are their diet (but, it has been noted, always supplemented by the minuscule animal life found on the seaweed—fish fed with pure seaweed soon fall into a decline).